2104100 - Disclosure - Acquisitions link:presentationLink link:calculationLink link:definitionLink 2404402 - Disclosure - Acquisitions - Additional Information (Details) link:presentationLink link:calculationLink link:definitionLink 2108100 - Disclosure - Commitments and Contingencies link:presentationLink link:calculationLink link:definitionLink 2408401 - Disclosure - Commitments and Contingencies - Additional Information (Details) link:presentationLink link:calculationLink link:definitionLink 1001000 - Statement - Consolidated Balance Sheets link:presentationLink link:calculationLink link:definitionLink 1001501 - Statement - Consolidated Balance Sheets (Parenthetical) link:presentationLink link:calculationLink link:definitionLink 1004000 - Statement - Consolidated Statements of Cash Flows link:presentationLink link:calculationLink link:definitionLink 1002000 - Statement - Consolidated Statements of Operations and Comprehensive Loss link:presentationLink link:calculationLink link:definitionLink 1003000 - Statement - Consolidated Statements of Stockholders’ Equity link:presentationLink link:calculationLink link:definitionLink 0002000 - Document - Document and Entity Information link:presentationLink link:calculationLink link:definitionLink 2109100 - Disclosure - Grant Award link:presentationLink link:calculationLink link:definitionLink 2409401 - Disclosure - Grant Award - Additional Information (Details) link:presentationLink link:calculationLink link:definitionLink 2107100 - Disclosure - Leased Properties link:presentationLink link:calculationLink link:definitionLink 2407402 - Disclosure - Leased Properties - Additional Information (Details) link:presentationLink link:calculationLink link:definitionLink 2407403 - Disclosure - Leased Properties - Future Minimum Lease Payments (Details) link:presentationLink link:calculationLink link:definitionLink 2407403 - Disclosure - Leased Properties - Future Minimum Lease Payments (Details) link:presentationLink link:calculationLink link:definitionLink 2307301 - Disclosure - Leased Properties (Tables) link:presentationLink link:calculationLink link:definitionLink 2103100 - Disclosure - License and Collaboration Agreements link:presentationLink link:calculationLink link:definitionLink 2403401 - Disclosure - License and Collaboration Agreements - Additional Information (Details) link:presentationLink link:calculationLink link:definitionLink 2110100 - Disclosure - Net Loss Per Share link:presentationLink link:calculationLink link:definitionLink 2410402 - Disclosure - Net Loss Per Share - Potential Dilutive Outstanding Securities Excluded From Diluted Net Loss Per Common Share (Details) link:presentationLink link:calculationLink link:definitionLink 2310301 - Disclosure - Net Loss Per Share (Tables) link:presentationLink link:calculationLink link:definitionLink 2101100 - Disclosure - Organization and Description of the Business link:presentationLink link:calculationLink link:definitionLink 2401401 - Disclosure - Organization and Description of the Business - Additional Information (Details) link:presentationLink link:calculationLink link:definitionLink 2104100 - Disclosure - Stock-Based Compensation link:presentationLink link:calculationLink link:definitionLink 2404402 - Disclosure - Stock Based Compensation - Additional Information (Details) link:presentationLink link:calculationLink link:definitionLink 2404405 - Disclosure - Stock-Based Compensation - Summary of Assumptions Used in Black-Scholes Option-Pricing Model for Stock Options Granted to Employees and Directors (Details) link:presentationLink link:calculationLink link:definitionLink 2404404 - Disclosure - Stock-Based Compensation - Summary of Recognized Stock-Based Compensation Expense Related to Stock Options Granted to Employees and Directors (Details) link:presentationLink link:calculationLink link:definitionLink 2404403 - Disclosure - Stock-Based Compensation - Summary of Stock Option Activity for Amended 2014 Plan and Related Information (Details) link:presentationLink link:calculationLink link:definitionLink 2304301 - Disclosure - Stock-Based Compensation (Tables) link:presentationLink link:calculationLink link:definitionLink 2106100 - Disclosure - Stockholders' Equity link:presentationLink link:calculationLink link:definitionLink 2406402 - Disclosure - Stockholders' Equity - Additional Information (Details) link:presentationLink link:calculationLink link:definitionLink 2406403 - Disclosure - Stockholders' Equity - Schedule of Common Stock Warrants Outstanding (Details) link:presentationLink link:calculationLink link:definitionLink 2306301 - Disclosure - Stockholders' Equity (Tables) link:presentationLink link:calculationLink link:definitionLink 2111100 - Disclosure - Subsequent Events link:presentationLink link:calculationLink link:definitionLink 2411401 - Disclosure - Subsequent Events (Details) link:presentationLink link:calculationLink link:definitionLink 2102100 - Disclosure - Summary of Significant Accounting Policies link:presentationLink link:calculationLink link:definitionLink 2402404 - Disclosure - Summary of Significant Accounting Policies - Additional Information (Details) link:presentationLink link:calculationLink link:definitionLink 2402403 - Disclosure - Summary of Significant Accounting Policies - Financial Assets and Liabilities Remeasured on a Recurring Basis (Details) link:presentationLink link:calculationLink link:definitionLink 2202201 - Disclosure - Summary of Significant Accounting Policies (Policies) link:presentationLink link:calculationLink link:definitionLink 2302302 - Disclosure - Summary of Significant Accounting Policies (Tables) link:presentationLink link:calculationLink link:definitionLink